Complicated and also quite different from the TeX ligkern model. Also let us not forget that OpenType is aimed for a GUI where you can activate and deactivate properties, and see what happens. The result of a bunch of combined OpenType transformations is not always trivial to predict, but when you are on a GUI you don't care about that, you just see what happens.
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...pour distinguer l'extérieur d'un aquarium,
mieux vaut n'être pas poisson
...the ball I threw while playing in the park
has not yet reached the ground
Es gab eine Zeit, wo ich nur ungern über Schubert sprechen,
nur Nächtens den Bäumen und Sternen von ihm vorerzählen mögen.